Vegetation of Nafud Al-Urayq
The terrain of Nafud Al-Urayq is located in an extra arid zone and
composed mainly of xeromorphic shrubs and herbs. Although the terrain
falls within a arid phytogeographical zone, there are intrusions of
species from other phytogeographical zones such as Acacias and
Astragalus of Somalia Masai and Irano-Turanian zones respectively.
Majority of species belonging to the Saharo-Arabian zone are annuals
with either xeromorphic adaptations or live only for a short period. The
duration of life of annuals in this area varies from year to year,
depending upon the favourable conditions. If the area receives
sufficient quantity of rain in a season, density and abundance of
herbaceous flora are significantly high while in years of drought, only
a few disseminules of the herbs could germinate and produce few flowers
and seeds. Depending on the amount precipitation, the overall stature of
these plants also varies from a retarded height to normal heights.
Haloxylon persicum is the major component of Nafud Al-Urayq,
dominating the northern half of the surveyed area, while Acacias (A.
ehrenbergiana, A. raddiana and A. tortilis) and xeromorphic
dwarf shrublets (Anvillea garcinii, Blepharis ciliaris, Gymnocarpos
decander, Sclerocephalus arabicus, etc. dominate the hillocks and
shallow depressions of the southern part. H. persicum is
characteristic of the mobile dune area which is also home to a number of
shrubs, subshrubs and herbs such as H. salicornicum, Moltkiopsis
ciliata, Schismus barbatus, Plantago boissieri, etc. Calligonum
comosum, a previously dominant population sharing with H.
persicum in the northern and central part is now in a highly
degraded state, majority of which are either dead or with a few live
branches. Herbaceous flora and other grasses are usually found in
depressions where silt and soil accumulates. Subshrubs and shrubs
dominate the southern edge of the protected area, mostly found at foot
hills or in crevices among hillocks. Haloxylon salicornicum
community is mainly dominating in the southern parts and along the
periphery of the northern half where the sand is less mobile. Non
palatable tropical element, Calotropis procera and the common
desert component Rhazya stricta are not prominent in the surveyed
area, primarily due to the restrictions to the domesticated herbs. The
palatable grass, Panicum turgidum dominates along the southern
side, where it shares the habitat of A. ehrenbergiana, and A.
tortilis and a number of annuals, such as Plantago spp.,
Brassica tourneforte, Centaurea pseudosinaica, Ifloga spicata,
Anisosciadium lanatum, etc. 23 locations were selected as sample
spots for recording the cover value of populations. An analysis of these
values has shown that, the entire vegetation can be divided into 5
communities or vegetation groups, each located in specific habitat. Some
species are widely distributed with appreciable abundance while others
are with restricted distribution range. The characteristic communities
are as follows:
Haloxylon salicornicum
community:
This community is highly scattered in the northern and southern part of
Nafud Al-Urayq.
It also dominates in the depressions and semi sabkha areas where rain
water accumulates during rainy days. Other species equally
dominant in this community are Pulicaria undulata and the
non-palatable species Euphorbia retusa (Fig. 6) and E.
dracunculoides. Major components of this community are:
Ajuga arabica,
Andrachne telephoides,
Capparis spinosa,
Citrullus colocynthis,
Euphorbia retusa,
E. dracunculoides,
Haloxylon salicornicum,
Pulicaria undulata
Haloxylon salicornicum-Pulicaria undulata
community:
dominates in the southern and northern edge of Nafud Al-Urayq, mostly
composed of herbaceous flora along the periphery and edges of sand dunes. This community is usually found on low altitude sand dunes of
southern sides and on sand sheets of the northern side with less mobile
sand. It contains the highest diversity both in terms of abundance and
density.
Presence of Capparis spinosa along with Pulicaria undulata
as a co-dominant species indicates the area as a shallow depression
receiving excess quantity of silt and sand.
Majority of species in this community other than the dominant ones are
herbs except for a species such as Kohautia caespitose, Senna italica,
etc. Low lying areas between dunes support a small population of A.
ehrenbergiana. Major components of this community are:
Acacia ehrenbergiana,
Anisosciadium lanatum,
Anthemis deserti,
Arnebia hispidissima,
Asphodelus tenuifolius,
Atractylis carduus,
Bassia muricata,
Capparis spinosa,
Cleome amblyocarpa,
Eremobium lineare,
Erodium laciniatum,
Fagonia glutinosa,
Farsetia stylosa,
Horwoodia dicksoniae,
Koelpinia linearis,
Kohautia caespitosa,
Launaea capitata,
L. mucronata,
Lotononis platycarpa,
Malva parviflora,
Neurada procumbens,
Pergularia tomentosa,
Plantago amplexicaulis,
P. boissieri,
P. ciliata,
Pulicaria undulata,
Reseda arabica,
Savignya parviflora,
Senna italica
Sabkha sub-community:
This community is the smallest among all other prominent ones and
therefore not significant in the study area.
Distribution of this community is highly restricted, found only along
the edges of water logged area.
It is dominated by
Tetraena propinqua associated with Haloxylon salicornicum.
Other associated species in the community are:
Cistanche phylepaea,
Haloxylon salicornicum,
Tamarix nilotica,
Tetraena propinqua (=Zygophyllum migahidii)
|
Species found in Nafud Al-Urayq
Population Cover:
A=Abundant; L=Locally Abundant; R=Rare
Chorotype:
SA= Saharo Arabian; SM= Somalia Masai; TR=Tropical; Med-SA=
Mediterranean –Saharo Arabian; Paleotropic; SA-IT= Saharo Arabia-Irano
Turanian; SA-SH= Saharo Arabia-Sahel; SA-SM= Saharo Arabian – Somalia
Masai; SM-SH=Somalia Masai-Sahel
Distribution:
Najd= Central Region; E= Eastern Province; N=Northern Region;
SW=Southwestern Region; NW=Northwestern Region; S=Southern Region.
Family |
Species |
Lifeform |
Population Cover |
Habitat |
Chorotype |
Distribution in Saudi Arabia |
Acanthaceae |
Blepharis ciliaris |
Herb |
L |
Foothills and Slopes |
SA-SH |
Najd, E, N, SW, S |
Amaranthaceae |
Aerva javanica |
Subshrub/ |
L |
Slopes of Hillocks |
Paleotropic |
Najd, E, N, SW, NW, S |
|
Bassia muricata |
Herb |
L |
Foothills |
SA-IT |
Najd, E, N, |
|
Halothamnus iraqensis |
Subshrub |
R |
Slopes of hills |
SA |
Najd, E, N |
|
Haloxylon persicum |
Shrub |
A |
Sand dunes |
IT |
Najd, E, N |
|
Haloxylon salicornicum |
Subshrub |
A |
Sand sheets |
SA |
Najd, N, E, SW, NW |
Apiaceae |
Anisosciadium lanatum |
Herb |
L |
Foothills and Slopes |
SA |
Najd, N, E, NW |
Apocynaceae |
Calotropis procera |
Shrub |
L |
Depressions, open plains |
SM-SH |
Najd, E, N, SW, NW, S |
|
Glossonema boveanum
ssp. boveanum |
Subshrub |
R |
Foothills and Slopes |
SM-SH |
Najd, N, NW |
|
Glossonema variens |
Subshrub |
R |
Foothills and Slopes |
SM-SH |
Najd, E, N, NW |
|
Pergularia tomentosa |
Subshrub |
R |
Foothills and Slopes |
SM-SH |
Najd, E, N, S |
Asphodelaceae |
Asphodelus refractus |
Herb |
R |
Depressions |
SA |
Najd, E, N, SW, NW, S |
|
Asphodelus tenuifolius |
Herb |
A |
Depressions |
SA-SH |
Najd, E, N, SW, NW, S |
Asteraceae |
Anthemis deserti |
Herb |
L |
Depressions |
SA |
Najd, E, N, NW |
|
Anvillea garcinii |
Subshrub |
L |
Foothills and Slopes |
SA |
Najd, E, N, NW |
|
Asteriscus heirochunticus |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N, S |
|
Atractylis carduus |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N |
|
Centaurea bruguieriana |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N |
|
Centaurea pseudosinaica |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N, SW, NW, S |
|
Centaurea sp. |
Herb |
|
Foothills and Slopes |
SA |
Najd, E, S |
|
Echinops polyceras |
Subshrub |
R |
Foothills and Slopes |
IT |
Najd, N |
|
Filago desertorum |
Herb |
A |
Depressions |
SA-IT |
Najd, E, N |
|
Ifloga spicata |
Herb |
A |
Foot hills, hill slopes, Depressions |
SA |
Najd, E, N |
|
Koelpinia linearis |
Herb |
L |
Foothills and Slopes |
SA-IT |
Najd, E, N |
|
Launaea capitata |
Herb |
A |
Open plains, depressions |
SA |
Najd, E, N, SW, NW, S |
|
Launaea mucronata |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N, SW, NW, S |
|
Launaea nudicaulis |
Herb |
L |
Foothills and Slopes |
SA |
Najd, E, N, SW, NW, S |
|
Picris babylonica |
Herb |
A |
Foothills and Slopes |
SA-IT |
Najd, E, N |
|
Pulicaria incisa |
Subshrub |
R |
Foothills and Slopes |
SA |
Najd, E, N, NW |
|
Pulicaria undulata |
Subshrub |
A |
Open plains |
SA-SM |
Najd, E, N, SW, NW, S |
|
Rhanterium epapposum |
Subshrub |
L |
Open plains |
SA |
Najd, E, N, NW |
|
Senecio flavus |
Herb |
L |
Slopes of Hillocks |
SA |
Najd, E, N, NW |
Boraginaceae |
Anchusa
sp. |
Herb |
R |
Slopes of Hillocks |
SA |
Najd, E, N, SW, NW |
|
Arnebia hispidissima |
Herb |
A |
Open plains |
SA |
Najd, E, N, SW, NW, S |
|
Heliotropium bacciferum |
Subshrub |
L |
Open plains |
SA-SH |
Najd, E, N, SW, NW, S |
|
Heliotropium digynum |
Subshrub |
A |
Sand dunes |
SA |
Najd, E, N, NW |
|
Moltkiopsis ciliata |
Subshrub |
A |
Sand dunes |
SA |
Najd, E, N, NW, S |
|
Trichodesma africanum |
Subshrub |
L |
Slopes of Hillocks |
SA-SH |
Najd, E, N, SW, NW, S |
Brassicaceae |
Brassica tourneforte |
Herb |
R |
Slopes of Hillocks |
Med-SA |
Najd, E, N |
|
Diplotaxis acris |
Herb |
R |
Slopes of Hillocks |
SA |
Najd, E |
|
Diplotaxis harra |
Herb |
R |
Slopes of Hillocks |
SA |
Najd, E |
|
Eremobium lineare |
Herb |
A |
Sand dunes |
SA |
Najd, E, N, NW |
|
Farsetia aegyptiaca |
Subshrub |
L |
Slopes of Hillocks |
SA-SH |
Najd, E, N, SW, NW, S |
|
Farsetia stylosa |
Subshrub |
L |
Slopes of Hillocks |
SA-SH |
Najd, E, N, SW, NW, S |
|
Horwoodia dcksoniae |
Herb |
L |
Foot hills |
SA |
Najd, E, N |
|
Morettia parviflora |
Subshrub |
L |
Slopes of Hillocks |
SM |
Najd, E, N |
|
Savignya parviflora |
Herb |
L |
Foot hills and depressions |
SA |
Najd, E, N |
|
Schimpera arabica |
Herb |
L |
Slopes of Hillocks |
SA |
Najd, E, N, SW, NW, S |
|
Zilla spinosa |
Herb |
L |
Foot hills |
SA |
Najd, E, N, SW, NW |
Capparaceae |
Capparis spinosa |
Shrub |
A |
Depressions |
Med-SA |
Najd, E, N |
|
Cleome amblyocarpa |
Herb |
L |
Depressions |
SA-SM |
Najd, E, N, SW, NW, S |
Caryophyllaceae |
Gymnocarpos decander |
Subshrub |
L |
Slopes of Hillocks |
SA |
Najd, E, N, NW |
|
Paronychia arabica |
Herb |
L |
Depressions |
SA |
Najd, E, N |
|
Polycarpaea repens |
Subshrub |
A |
Sand dunes |
SA-SM |
Najd, E, N, NW |
|
Sclerocephalus arabicus |
Subshrub |
L |
Slopes of Hillocks |
SA |
Najd, E, N, NW, S |
|
Silene villosa |
Herb |
A |
Sand dunes |
SA |
Najd, E, N, NW |
|
Spergula fallax |
Herb |
L |
Depressions |
SA |
Najd, E, N, NW, S |
Cistaceae |
Helianthemum lippii |
Subshrub |
R |
Slopes of Hillocks |
SA-SM |
Najd, E, N, NW |
Convolvulaceae |
Convolvulus buschiricus |
Subshrub |
R |
Slopes of Hillocks |
SA |
Najd, E, N, NW, S |
|
Convolvulus oxyphyllus
ssp. oxycladus |
Subshrub |
R |
Open plains |
SA |
Najd, E, N, NW, S |
|
Convolvulus pilosellifolius |
Subshrub |
L |
Depressions |
IT |
Najd, E, N, NW, S |
Cucurbitaceae |
Citrullus colocynthis |
Subshrub |
A |
Depressions |
SA |
Najd, E, N, SW, NW, S |
Cynomoriaceae |
Cynomorium coccineum |
Subshrub |
R |
Sand dunes |
Med-IT-SA |
Najd, E, N |
Cyperaceae |
Cyperus conglomeratus |
Subshrub |
A |
Sand dunes |
SA |
Najd, E, N, NW |
Ephedraceae |
Ephedra foliata |
Shrub |
R |
Slopes of Hillocks |
SA-IT |
Najd, E, N, SW, NW, S |
Euphorbiaceae |
Andrachne telephioides |
Subshrub |
L |
Depressions |
Med-IT |
Najd, E, N |
|
Euphorbia dracunculoides |
Subshrub |
A |
Depressions |
SA |
Najd, E |
|
Euphorbia granulata |
Subshrub |
A |
Open plains, depressions |
SA-SM |
Najd, E, N, SW, NW, S |
|
Euphorbia retusa |
Subshrub |
A |
Depressions |
SA |
Najd, E, N, NW |
Fabaceae |
Acacia raddiana |
Tree |
A |
Open plains, wadi channels |
SM |
Najd, E, N |
|
Acacia ehrenbergiana |
Shrub |
L |
Open plains, wadi channels |
SM |
Najd, E, N, NW |
|
Acacia tortilis |
Shrub |
L |
Open plains, wadi channels |
SM |
Najd, E, N, SW, NW, S |
|
Astragalus caprinus |
Herb |
R |
Depressions |
SA-IT |
Najd, E, N |
|
Astragalus schimperi |
Herb |
L |
Depressions |
SA-IT |
Najd, E, N |
|
Astragalus tribuloides |
Herb |
L |
Depressions |
SA-IT |
Najd, E, N, NW |
|
Astragalus spinosus |
Subshrub |
L |
Depressions |
SA-IT |
Najd, E, N |
|
Lotononis platycarpos |
Herb |
A |
Open plains, Sand sheets |
SA-SM |
Najd, E, N, NW |
|
Medicago laciniata |
Herb |
L |
Depressions, slopes of hillocks |
SA |
Najd, E, N, NW, S |
|
Senna italica |
Subshrub |
L |
Open plains |
SM |
Najd, E, N, SW, NW, S |
|
Tephrosia purpurea |
Subshrub |
R |
Open plains
Slopes of hills |
SM |
Najd, E, N, SW, NW, S |
Geraniaceae |
Erodium laciniatum |
Herb |
L |
Open plains |
Med |
Najd, E, N, SW, NW, S |
|
Monsonia heliotropoides |
Herb |
R |
Open plains |
SM |
Najd, E, NW |
|
Monsonia nivea |
Herb |
A |
Sand dunes |
SA-SM |
Najd, E, N |
Lamiaceae |
Ajuga arabica |
Subshrub |
L |
Depressions |
IT |
Najd, N |
|
Salvia deserti |
Subshrub |
R |
Slopes of hills |
SA |
Najd, E, N, NW, S |
|
Teucrium polium |
Subshrub |
R |
Slopes of hills |
Med-IT |
Najd, E, N, NW |
Liliaceae |
Gagea
sp. |
Herb |
R |
Open plains |
Med-SA |
Najd, E, N |
Malvaceae |
Althaea ludwigii |
Herb |
L |
Depressions |
SA |
Najd, E, N, SW, NW, S |
|
Hibiscus micranthus |
Shrub |
L |
Slopes of hills |
Paleotropic |
Najd, E, N, SW, NW, S |
|
Malva parviflora |
Herb |
L |
Depressions |
Med-IT |
Najd, E, N, SW, NW, S |
Molluginaceae |
Gisekia pharnaceoides |
Herb |
A |
Open plains, sand sheets |
TR |
Najd, E, NW, S |
Neuradaceae |
Neurada procumbens |
Herb |
A |
Sand sheets |
SA |
Najd, E, N, NW |
Orobanchaceae |
Cistanche phelypaea |
Subshrub |
L |
Sand dunes |
SA-IT |
Najd, E, N |
Plantaginaceae |
Plantago amplexicaulis |
Herb |
L |
Slopes of hills |
SA |
Najd, E, N, SW, NW, S |
|
Plantago boissieri |
Herb |
A |
Sand dunes, sand sheets |
SA |
Najd, E, N, NW |
|
Plantago cilata |
Herb |
A |
Depressions and sand sheets |
SA |
Najd, E, N |
|
Plantago ovata |
Herb |
A |
Depressions and sand sheets |
SA-IT |
Najd, E, N |
Poaceae |
Aristida adscensionis |
Annual |
A |
Sand dunes |
SA |
Najd, E, N, NW |
|
Aristida funiculata |
Perennial |
L |
Sand dunes |
SA |
Najd, E, N |
|
Brachypodium distachyon |
Annual grass |
L |
Slopes of hills |
Med-IT-SA |
Najd, E, N, NW, S |
|
Cenchrus ciliaris |
Perennial grass |
L |
Slopes of hills |
TR |
Najd, E, N, SW, NW, S |
|
Centropodia forsskalii |
Perennial |
A |
Sand dunes |
SA-IT |
Najd, E, N |
|
Chrysopogon plumulosus |
Perennial |
R |
Slopes of hills |
SA-SM |
Najd, E, N, NW, S |
|
Cynodon dactylon |
Perennial |
A |
Slopes of hills, depressions |
TR |
Najd, E, N, SW, NW, S |
|
Enneapogon desvauxii |
Perennial |
L |
Slopes of hills |
SA-IT |
Najd, E, N |
|
Eragrostis barrelieri |
Perennial |
L |
Slopes of hills |
Med-SA |
Najd, E, N, SW, NW, S |
|
Hyparrhenia hirta |
Annual |
L |
Slopes of hills |
Med-IT-SA |
Najd, E, N |
|
Lolium persicum |
Annual |
R |
Slopes of hills, depressions |
IT |
Najd, E, N, SW, NW, S |
|
Ochthochloa compressa |
Perennial |
L |
Slopes of hills |
SA-IT |
Najd, E |
|
Panicum turgidum |
Perennial
|
A |
Open plains, foot hills |
SA-SM |
Najd, E, N, SW, NW, S |
|
Phalaris minor |
Annual |
L |
Depressions |
Med-IT |
Najd, E, N, NW |
|
Schismus barbatus |
Annual |
L |
Depressions |
SA-IT |
Najd, E, N |
|
Stipagrostis ciliata |
Perennial |
R |
Sand dunes |
SA |
Najd, E, N, NW |
|
Stipagrostis drarii |
Perennial |
A |
Sand dunes |
SA |
Najd, E, N, NW, S |
|
Stipagrostis hirtigluma |
Perennial |
R |
Sand dunes |
SA |
|
|
Stipagrostis obtusa |
Perennial |
L |
Sand dunes |
SA-IT |
Najd, E, N |
|
Stipagrostis plumosa |
Perennial |
A |
Sand dunes |
SA-IT |
Najd, E, N |
|
Tetrapogon villosus |
Perennial |
L |
Slopes of hills |
SA-SM |
Najd, E |
Polygalaceae |
Polygala erioptera. |
Subshrub |
R |
Slopes of hills |
SA-SM |
Najd, E, N, SW, NW |
Polygonaceae |
Calligonum comosum |
Shrub |
L |
Sand dunes |
SA-IT |
Najd, E, N |
|
Emex spinosa |
Herb |
L |
Depressions |
Med-SA |
Najd, E, N, SW, NW, S |
|
Rumex vesicarius |
Herb |
L |
Slopes of hills, Depressions |
SA |
Najd, E, N, SW, NW, S |
Polypodiaceae |
Cheilanthus sp. |
Subshrub |
R |
Moist hill slopes |
SA-IT |
Najd, E, N, SW, NW |
Resedaceae |
Ochradenus baccatus |
Shrub |
L |
Slopes of hills |
SA-SM |
Najd, E, N, SW, NW, S |
|
Reseda arabica |
Herb |
R |
Depressions |
SA |
Najd, E |
Rubiaceae |
Kohautia caespitosa |
Subshrub |
R |
Slopes of hills |
SA-SM |
Najd, E, N, SW, NW, S |
Rutaceae |
Haplophyllum tuberculatum |
Subshrub |
L |
Open plains |
SA |
Najd, E |
Solanaceae |
Lycum shawii |
Shrub |
L |
Foot hills |
SA-SH |
Najd, E, N, SW, NW, S |
Tamaricaceae |
Tamarix aphylla |
Tree |
R |
Abandoned farms |
SM |
Najd, E, N, SW, NW, S |
|
Tamarix nilotica |
Shrub |
L |
Saline area/sabkha |
SA |
Najd, E, N, SW, NW, S |
Urticaceae |
Forsskaolea tenacissima |
Subshrub |
L |
Slopes of hills |
SA-SM |
Najd, E, N, SW, NW, S |
Zygophyllaceae |
Fagonia bruguieri |
Subshrub |
A |
Slopes of hills |
SA |
Najd, E, N, SW, NW, S |
|
Fagonia glutinosa |
Subshrub |
A |
Sand sheets |
SA |
Najd, E, N, NW, S |
|
Seetzenia lanata |
Herb |
L |
Sand sheets |
SM |
Najd, E |
|
Tetraena propinqua |
Subshrub |
A |
Saline area/sabkha |
SA |
Najd, E, N, NW |
|
Tribulus terrestris
var. terrestris |
Herb |
L |
Depressions, hill slopes |
Paleotropic |
Najd, E, N, SW, NW, S |
|